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ildings; although here a form of social segregation did prevail, whereby shopkeepers and tradesmen tended to occupy the cheaper-to-rent cellars and garrets, while the more well-to-do professional classes occupied the more expensive middle storeys. During the Jacobite rising of 1745, Edinburgh was briefly occupied by the Jacobite "Highland Army" before its march into England. After its eventual defeat at Culloden, there followed a period of reprisals and pacification, largely directed at the rebellious clans. In Edinburgh, the Town Council, keen to emulate London by initiating city improvements and expansion to the north of the castle, reaffirmed its belief in the Union and loyalty to the Hanoverian monarch George III by its choice of names for the streets of the New Town: for example, Rose Street and Thistle Street; and for the royal family, George Street, Queen Street, Hanover Street, Frederick Street and Princes Street (in honour of George's two sons). In the second half of the century, the city was at the heart of the Scottish Enlightenment, when thinkers like David Hume, Adam Smith, James Hutton and Joseph Black were familiar figures in its streets. Edinburgh became a major intellectual centre, earning it the nickname "Athens of the North" because of its many neo-classical buildings and reputation for learning, recalling ancient Athens. In the 18th-century novel The Expedition of Humphry Clinker by Tobias Smollett one character describes Edinburgh as a "hotbed of genius". Edinburgh was also a major centre for the Scottish book trade. The highly successful London bookseller Andrew Millar was apprenticed there to James McEuen. From the 1770s onwards, the professional and business classes gradually deserted the Old Town in favour of the more elegant "one-family" residences of the New Town, a migration that changed the city's social character. According to the foremost hi storian of this development, "Unity of social feeling was one of the most valuable heritages of old Edinburgh, and its disappearance was widely and properly lame